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Publications by Richard L. Brown


Brown, R.L. 1974.  Diets and habitat preferences of selected anurans in Southeast Arkansas. Amer. Midl. Nat. 91: 468-473.

Brown, R.L. & R.T. Allen. 1974.  Larval foodplants and parasites of some Lepidoptera in Southeast Arkansas.  Journ. Lepid. Soc. 28: 168-170.

Brown, R.L. 1976.  Behavioral observations on Aethalion reticulatum (Hem., Aethalionidae) and associated ants.  Insectes Sociaux 23: 99-108.

Brown, R.L. 1979.  Nomenclatorial changes in Eucosmini (Tortricidae). Journ. Lepid. Soc. 33: 21-28.

Brown, R.L. 1979.  The valid generic and tribal names for the codling moth, Cydia pomonella (Olethreutinae: Tortricidae).  Ann. Entomol. Soc. Amer. 72: 565-567.

Brown, R.L. 1980.  A new species of Epinotia Hubner (Lepidoptera: Tortricidae). Proc. Entomol. Soc. Washington 82: 504-509.

Brown, R.L. 1980. A correction of the name for the type-species of Rhopobota Lederer (Tortricidae: Eucosmini). Journ. Lepid. Soc. 34: 325.

Brown, R.L. 1982.  Notes on Gretchena: a new species and the synonymy of Gwendolina (Lepidoptera: Tortricidae). Proc. Entomol. Soc. Washington 84: 594-602.

Roelofs, W. & R.L. Brown. 1982.  Pheromones and evolutionary relationships of Tortricidae.  Ann. Rev. Ecol. Syst. 13: 395-422.

Brown, R.L. & W.E. Miller. 1983.  Valid names of the spruce seed moth and a related Cydia species (Lepidoptera: Tortricidae).  Ann. Entomol. Soc. Amer. 76: 110-111.

Brown, R.L. 1984.  Taxonomic and morphological investigations of Olethreutinae:  Rhopobota, Griselda, Melissopus, and Cydia (Lepidoptera: Tortricidae). Entomography 2: 97-120.

Brown, R.L. and P.R. Miller. 1984.  Studies of Lepidoptera hindwings with emphasis on ultrastructure of scales of Cydia caryana (Fitch) (Tortricidae). Entomography 2: 261-295.

Brown, R.L. 1984.  Review of Corticivora (Lepidoptera: Tortricidae) with descriptions of new species and analysis of their tribal relationships. Proc. Entomol. Soc. Washington 86: 278-286.

Brown, R.L., J.F.G. Clarke, and D.H. Habeck. 1984.  New host records for Olethreutinae (Tortricidae). Journ. Lepid. Soc.  37: 224-227.

Brown, R.L. 1987. Tortricoidea. pp. 419-433. In Immature Insects, Ed. F. Stehr. Kendall/Hunt Publ. Co. 754 pp.

Adamski, D. and R.L. Brown. 1987.  A new Nearctic Glyphidocera with descriptions of all stages (Lepidoptera: Blastobasidae: Symmocinae). Proc. Entomol. Soc. Washington 89: 329-343.

Brown, R.L. 1987. Ressurection of Catastega Clemens and revision of the Epinotia vertumnana species-group (Tortricidae: Olethreutinae). Journ. Lepid. Soc. 40: 327-346.

Adamski, D. & Brown, R.L.  1989. Morphology and systematics of North American Blastobasidae [Lepidoptera: Gelechioidea].  Miss. Agric. For. Exp. Sta. Tech. Bull. 165 (MEM no. 1): 1-70.

Horak, M. & R.L. Brown. 1991. Taxonomy and phylogeny. pp. 23-50.  In. L.P.S. van der Geest and H.H. Evenhuis (Eds.).  Tortricoid Pests. Their Biology, Natural Enemies and Control.  Elsevier Science Publ. 808 pp.

Brown, R.L. & J.A. Powell. 1991. A new species of Epiblema (Tortricidae: Olethreutinae) from coastal redwood forests in California with an analysis of the forewing pattern.  Pan-Pacific Entomol. 67: 107-114.

Allen, R.T. & R.L. Brown.  1991. The biota of Magazine Mountain (II): a preliminary list of the macrolepidoptera fauna.  Arkansas Acad. Sci. 45: 18-21.

Brown, R.L. 1992. Six new species of Catastega (Lepidoptera: Tortricidae: Eucosmini) from Mexico and Southwestern United States. J. New York Entomol. Soc. 100: 209-227.

Baker, G.T. &  R.L. Brown.  1995. Chorionic fine structure of the egg of the oak tingid, Corythuca arcuata (Say) (Hemiptera: Tingidae). Proc. Entomol. Soc. Washington 96: 70-73.

Powell, J.A., J. Razowski, & R.L. Brown.  1995. Olethreutinae.  pp. 151-157. In J.B. Heppner (Ed.). Atlas of Neotropical Lepidoptera. Checklist, Part 2.  W. Junk Publ.

Wright, D.J., R.L. Brown, & L.D. Gibson. 1997.  A new species of Phaneta, with taxonomic diagnoses and seasonal and geographical data on four related species (Tortricidae).  J. Lepid. Soc.   51: 119-127.

Cordero, R., R.L. Brown, and H.P. Pitre. 2000. Description of Life Stages and distribution of Metaponpneumata rogenhofferi (Lepidoptera: Noctuidae). Tropical Lepidoptera 10: 59-67.

Hickling, R. and R.L. Brown.  2000.  Analysis of acoustic communication by ants. Journ. Acoustical Soc. Amer. 108: 1920-1929.

Hickling, R. and R.L. Brown.  2001.  Response to "Ants are deaf."  Journ. Acoustical Soc. Amer. 109: 3083.

Adamski, D. and R. L. Brown.  2001.  Glyphidocera Walsingham (Lepidoptera:  Gelechioidea: Glyphidoceridae) of de la Neblina and adjacent areas in Amazonas, Venezuela.  Proc. Entomol. Soc. Washington 103: 968-998.

Miller, W. E., R.L. Brown, and K. Tuck.  2001. Taxonomic clarification of Notocelia rosaecolana (Doubleday) and N. trimaculana (Haworth) (Tortricidae).  Journ. Lepid. Soc. 54: 111-118.

Adamski, D.  and R. L. Brown. 2002.  New species of Blastobasis Zeller from New Caledonia and Fiji (Lepidoptera: Gelechioidea: Coleophoridae: Blastobasinae).  Ins. Koreana 19: 137-145.

Brown, R.L. 2003. Paleoenvironment and biogeography of  the Mississippi Black Belt.  Evidence from insects.  pp. 11-26.  In E. Peacock and T. Schauwecker (eds.). Blackland Prairies of the Gulf Coastal Plain: Culture, Nature and Sustainability. Univ. Alabama Pr. 348 pp.

Wiygul, S., Krans, K., Brown, R., and Maddox. 2003. Restoration of a prairie remnant in the Black Belt of Mississippi.   pp. 254-261. In E. Peacock and T. Schauwecker (eds.).  Blackland Prairies of the Gulf Coastal Plain: Nature, Culture, and Sustainability.  Univ. Alabama Pr. 348 pp.

Wagner, D.L., Adamski, D.,  and Brown, R.L.  2004.  A  species of Mompha Hubner (Lepidoptera: Coleophoridae: Mompinae) from buttonbush (Cephalanthus occidentalis L.) with descriptions of the early stages.  Proc. Entomol. Soc. Washington 106: 1-18.

Brown, J.W. and Brown, R.L.  2004. A new species of Cryptaspasma (Lepidoptera: Tortricidae: Olethreutinae) from Central America, the Caribbean, and Southeastern United States, with a catalog of the world fauna of Microcorsini.  Proc. Entomol. Soc. Washington 106:  288-297...

Vogt, J.T., Reed, J.T. and Brown, R.L. 2004.  Temporal foraging activity of selected ant species in northern Mississippi during summer months.  J. Entomol. Sci. 39:  444-452.

Baixeras, J., Brown, J., Brown, R., Horak, M., Komai, F., Metzler, E., Razowski, J., Tuck, K. 2005.  In Brown, J. (Ed.).  World Catalog of Insects. Vol. 5.  Tortricidae (Lepidoptera).  Apollo Books.  608 pp.

Hill, J. and Brown, R.L.  2005.  The first record of the genus Polyergus (Hymenoptera: Formicidae) in Mississippi.  J. Miss. Acad. Sci.  50: 120.

Pinkaew, N, Chandrapatya, A.,  and Brown, R.L.  2005.  Two new species and a new record of Eucoenogenes (Meyrick) (Lepidoptera: Tortricidae) from Thailand with a discussion of characters defining the genus. Proc. Entomol. Soc. Washington 107:  869-882.

Landry, B. and Brown, R.L.  Two new species of Neodactria Landry Lepidoptera: Pyralidae, Crambinae) from the United States of America.  Zootaxa 1081: 1-16.

MacGown, J., Brown, R.L., and Hill, J. 2005 An annotated list of the Pyramica (Hymenoptera:  Formicidae: Dacetini) of Mississippi.  J.  Kansas Entomol. Soc. 78: 285-289.

MacGown, J.  and Brown, R.L. 2006. Survey of ants (Hymenoptera:  Formicidae) of the Tombigbee National Forest in Mississippi.  J. Kansas Entomol. Soc. 79: 325-340.

Hill, J. G., R. L. Brown, and J. A. MacGown. 2006. Environmental variables affecting ant (Formicidae) community composition in Mississippi’s Black Belt Prairie and Flatwoods regions.  Journal of the Mississippi Academy of Sciences 51: 56.

Lee, S. and R.L. Brown.  2006.  Descriptions of males of two species of Dichomeris (Lepidoptera: Gelechiidae) species with new distributional records in Mississippi and Alabama.  J. Lepid. Soc. 60: 38–40.

Lee, S.M. and R.L. Brown.  2006.  A new method for preparing slide mounts of whole bodies of microlepidoptera.  Journ. Asia–Pacific Entomol. 9: 249–253.

MacGown, J. A., R. L. Brown, and J. G. Hill. 2006. Carpenter ants (Camponotus) of Mississippi. Journal of the Mississippi Academy of Sciences 51: 115.

MacGown, J. A., R. L. Brown, J. G. Hill, and B. Layton. 2007. Carpenter Ants (Hymenoptera: Formicidae: Camponotus) of Mississippi. Mississippi Agricultural and Forestry Experiment Station Technical Bulletin 1158. 28 pp. [pdf]

Martinez, E. and R.L. Brown.  2007.  Argyrini (Lepidoptera: Crambidae) of Mississippi and Alabama with a redescription of Argyria rufisignella (Zeller). Journ. Lepid. Soc. 61:  78–83.

Brown, R.L. 2008.  Book review.  Olethreutine Moths of Australia (Lepidoptera: Tortricidae) – Marianne Horak. Syst. Entomol. 33: 212–214.

Hill, J.G., K.S. Summerville, and R.L. Brown.  2008.  Habitat associations of ant species Hymenoptera: Formicidae) in a heterogeneous Mississippi landscape.  Environ. Entomol. 37: 453–463.

Lee, S. and R.L. Brown.   2008.  Revision of Holarctic Teleiodini (Lepidoptera: Gelechiidae).  Zootaxa 1818: 1–55.

Lee, S.M. and R.L. Brown.   2008.  Phylogenetic Relationships of Holarctic Teleiodini Lepidoptera: Gelechiidae) Based on Analysis of Morphological and Molecular Data. Syst. Entomol. 33: 595–612.

Simonsen, T.J., R.L. Brown, and F.A.H. Sperling. 2008.  Tracing an invasion:  phylogeography of Cactoblastis cactorum (Lepidoptera: Pyralidae) in the United States based on mitochondrial DNA.  Ann. Entomol. Soc. Amer. 101: 899–905.

Lee, S., R.L. Brown and W. Monroe.  2009.  Use of confocal laser scanning microscopy in systematics of insects with a comparison of fluorescence from different stains. Syst. Entomol. 34: 10–14.

Lee, S., R.W. Hodges, and R.L. Brown. 2009.  Checklist of Gelechiidae (Lepidoptera) in American North of Mexico.  Zootaxa 2231: 1–39.

Siebert, M.W. and R.L. Brown. 2009.  Collection of Moodna bisinuella (Lepidoptera: Pyralidae) from field corn in Mississippi.  Midsouth Entomol. 2: 53–54.

Lee, S.M. and R.L. Brown. 2010. Review of Symmocinae (Lepidoptera:  Autostichidae), in North America with the description of a new genus and species. Journ. Lepid. Soc. 64:  177–187.

Hill, J. G. and R. L. Brown. 2010. The ant (Hymenoptera: Formicidae) fauna of Black Belt Prairie remnants in Alabama and Mississippi. Southeastern Naturalist 9: 73-84. [pdf]

Baker, G.T., S.D. Hight, and R.L. Brown.  2012.  External morphology of the egg of the native (Melitara prodenialis) and exotic (Cactobastis cactorum) cactus moths (Lepidoptera: Pyralidae).   Proc. Entomol. Soc. Washington 114: 433–438.

Lee, S. and R.L. Brown. 2012. A new species of Euclemensia (Cosmopterigidae) from the United States.  Journ. Lepid. Soc. 65: 47–50.

MacGown, J. A., J. G. Hill, T. L. Schiefer, and R. L. Brown.  2012. Ant diversity and habitat associations at the Noxubee National Wildlife Refuge in Mississippi. Mississippi Agricultural and Forestry Experiment Station Technical Bulletin 1197. [pdf]

Smith, B.A., R.L. Brown, W. LaBerge, and T. Griswold.  2012.  Faunistic study of bees (Hymenoptera: Apoidea) in the Black Belt Prairie of Mississippi.  Journ. Kansas Entomol. Soc. 85: 32–47. 

Lee, S. and R.L. Brown.  2013.  Pubitelphusa (Lepidoptera: Gelechiidae: Litini) —A new genus for two species assigned to Telphusa and Concubina. Proc. Entomol. Soc. Washington 115: 70–74.

MacGown, J., H. Richter, and R.L. Brown. 2013. Notes and new distribution records of invasive ants (Hymenoptera: Formicidae) in the southeastern United States. Midsouth Entomol.  6: 104-114.  Online at:

Seltzer, J., T. Schiefer, and R.L. Brown. 2013. Results from the Regional Identification Center of the USDA-APHIS (Eastern Region) at the Mississippi Entomological Museum for the 2012 and 2013 wood boring beetle surveys, including new state and county records for Alabama and Mississippi. Midsouth Entomol.  6: 119-133. Online at:

Brown, R.L. and C. Jaeger. 2014. A new species of Eucosma Hubner and two new species of Cydia Hubner (Lepidoptera: Tortricidae) from the United States.  Zootaxa 3860: 226-234.

Wright, D.J. and R.L. Brown.  2014. Review of the Eucosma refusana (Walker) species group with descriptions of two new species and discussion of biogeographic influences on species distributions.  Journal Lepidopterists Society 68: 85-100.

Schachat, S.R. and R.L. Brown. 2015. Color pattern on the forewing of Micropterix (Lepidoptera: Micropterigidae): insights into the evolution of wing pattern and wing venation in moths.  PLoS ONE 10(10): e0139972. doi: 10.1371/journal.pone.0139972.  

Schachat, S.R. and R.L. Brown. 2016. Forewing color pattern in Micropterigidae (Insecta: Lepidoptera): homologies between contrast boundaries, and a revised hypothesis for the origin of symmetry systems. BMC Evolutionary Biology 16:116: 1-25. [PDF]


Orkin Insect Zoo Entomology Curriculum Module (with Joe MacGown and MSU Center Education Technology and Development). 1998. URL:


Brown, R.L. 1989.  Evolution of aquatic adaptations in insects.  Proc. 35th Miss. Insect Control Conf. (abstract).

Brown, R.L. & S. Marshall. 1991. ECN Discussion Group Summary:  The role of small systematic institutions.  Insect Collection News 6: 3.

Brown, R.L. 1992. Looper complex and other Plusiinae larvae on soybean in southern United States: a seasonal profile (abstract).  Soybean Looper Resistance Workshop 1992 Presentation Abstracts.

Brown, R.L. 1995.  Lepidoptera in the Mississippi Entomological Museum.  Southern Lepidopterists' News 17 (2): 10-12.

Hickling,R., R.L. Brown, P. Lee, G. Stratton, and D. Velea.  1998.Analysis of acoustic communication by the Black Fire Ant Solenopsis richteri.  In:  Proceedings of the 1998 Imported Fire Ant Research Conference.  Hot Springs, AK  April 6-8 , 1998.  pp. 87-95. 

Brown, R.L and Martinez, E. 2004.  The Cactus Moth:  An Invading Pest.Information Brochure printed by MSU Printing Services.  Also available at following web site: